. 2016 Feb 13;16(1):203–229. doi: 10.1007/s12311-016-0763-3

Table 1.

Different learning rules in cerebral cortex, basal ganglia, and cerebellum

Brain site	Cerebral cortex	Basal ganglia	Cerebellar cortex
Cellular site	Excitatory afferents onto pyramidal cells	Striatum cortical afferents onto spiny neurons	Parallel fibers onto Purkinje cells
Learning Rule & operation	Hebbian & coincidence	Reinforcement & coincidence · reward	Supervised & coincidence · error
Permissive factors	Cholinergic +	Dopaminergic +	Noradrenergic +

This table illustrates key features of three learning rules in the brain (adapted with permission from Houk [113])