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. 1998 Oct 1;112(4):409–422. doi: 10.1085/jgp.112.4.409

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The sum total of times all flickering pores plus the subsequent successful pore were in open states before full pore enlargement. Total open times were the same for all symmetrical membranes, regardless of whether probe was present. For both R18 (left) and DiI (right), total open times were the greatest for probe accumulated in the cis monolayer and least for probe accumulated in the trans monolayer.